The Cenozoic is divided into three periods: The Paleogene, Neogene, and Quaternary; and seven epochs: The Paleocene, Eocene, Oligocene, Miocene, Pliocene, Pleistocene, and Holocene. The Quaternary Period was officially recognized by the International Commission on Stratigraphy in June 2009, and the former Tertiary Period was officially disused in 2004. The common use of epochs during the Cenozoic helps paleontologists better organize and group the many significant events that occurred during this comparatively short interval of time. There is also more detailed knowledge of this era than any other because of the relatively young strata associated with it.

Geologically, the Cenozoic is the era when the continents moved into their current positions. Australia-New Guinea, having split from Pangea during the early Cretaceous, drifted north and, eventually, collided with South-east Asia; Antarctica moved into its current position over the South Pole; the Atlantic Ocean widened and, later in the era, South America became attached to North America.

India collided with Asia 55 to 45 million years ago; Arabia collided with Eurasia, closing the Tethys ocean, around 35 million years ago.

**The concepts of Tertiary and Quaternary have an interesting history. In the 1760s and 1770s a geologist named Giovanni Arduino was studying the rocks and minerals in Tuscany. He classified mountains according to the type of rocks that he found in them. Unfossiliferous schists, granites, and basalts (all volcanic rocks) that formed the cores of large mountains he called Primitive. Fossil-rich rocks of limestone and clay that were found on the flanks of mountains over the Primitive rocks were called Secondary. Finally, there were another group of fossiliferous rocks of limestones and sandstones lying over the Secondary rocks and forming the foothills of the mountains that Anduino called Tertiary. So at first, Tertiary referred to a certain type of rock found in the area of Tuscany. But later, geologists used the fossils found in the Tertiary rocks there to recognize rocks of the same age elsewhere. Rocks with the same species of fossils were the same age.

Extensive Tertiary age rocks were recognized in the Paris Basin, which is the area around Paris, France. In the 1820s and 1830s Charles Lyell, a noted English geologist who had a great influence on Charles Darwin, subdivided the Tertiary rocks of the Paris Basin on their fossils. Lyell came up with an ingenious idea. He noticed that the rocks at the top of the section had a very high percentage of fossils of living mollusc species. Those at the bottom of the section had very few living forms. He deduced that this difference was because of the extinction of older forms and the evolution of living forms during the time that the rocks were being deposited. He divided the Tertiary rocks into three sub-ages: the Pliocene, the Miocene, and the Eocene. 90% of the fossil molluscs in Pliocene rocks were living today. In the Miocene rocks, only 18% of the molluscs were of living species, and in Eocene rocks, only 9.5%.

These subdivisions of the Tertiary have been correlated around the world using the fossil species in them. Rocks with the same species as Lyell’s Eocene, are considered to be the same age as those in the Paris Basin. The same goes for the other subdivisions. Some time later it was noted that in areas other than the Paris Basin, there were rocks that seemed to be from time periods that were not represented in Lyell’s sequence. This was because during those periods there had been no deposition in what would later be the Paris Basin. These two periods, later designated Oligocene and Paleocene, were inserted into the Tertiary in their proper places.

The Paleocene–Eocene Thermal Maximum of 55.8 million years ago was a significant global warming event. However, since the Azolla event of 49 million years ago, the Cenozoic Era has been a period of long-term cooling. After the tectonic creation of Drake Passage, when South America fully detached from Antarctica during the Oligocene, the climate cooled significantly due to the advent of the Antarctic Circumpolar Current which brought cool deep Antarctic water to the surface. The cooling trend continued in the Miocene, with relatively short warmer periods. When South America became attached to North America creating the Isthmus of Panama, the Arctic region cooled due to the strengthening of the Humboldt and Gulf Stream currents, eventually leading to the glaciations of the Quaternary ice age, the current interglacial of which is the Holocene Epoch.

During the Cenozoic, mammals proliferated from a few small, simple, generalized forms into a diverse collection of terrestrial, marine, and flying animals, giving this period its other name, the Age of Mammals, despite the fact that birds still outnumbered mammals two to one. The Cenozoic is just as much the age of savannas, the age of co-dependent flowering plants and insects, and the age of birds. Grass also played a very important role in this era, shaping the evolution of the birds and mammals that fed on it. One group that diversified significantly in the Cenozoic as well were the snakes. Evolving in the Cenozoic, the variety of snakes increased tremendously, resulting in many colubrids, following the evolution of their current primary prey source, the rodents.

In the earlier part of the Cenozoic, the world was dominated by the gastornid birds, terrestrial crocodiles like Pristichampsus, and a handful of primitive large mammal groups like uintatheres, mesonychids, and pantodonts. But as the forests began to recede and the climate began to cool, other mammals took over. The Cenozoic is full of mammals both strange and familiar, including chalicotheres, creodonts, whales, primates, entelodonts, saber-toothed cats, mastodons and mammoths, three-toed horses, giant rhinoceros like Indricotherium, and brontotheres.

Following the Paleozoic, the Mesozoic extended roughly 186 million years, from 252.2 to 66 million years ago when the Cenozoic Era began. This time frame is separated into three geologic periods. From oldest to youngest:

The lower (Triassic) boundary is set by the Permian-Triassic extinction event, during which approximately 90% to 96% of marine species and 70% of terrestrial vertebrates became extinct. It is also known as the “Great Dying” because it is considered the largest mass extinction in the Earth’s history. The upper (Cretaceous) boundary is set at the Cretaceous-Tertiary (KT) extinction event (now more accurately called the Cretaceous–Paleogene (or K–Pg) extinction event), which may have been caused by the impactor that created Chicxulub Crater on the Yucatán Peninsula. Towards the Late Cretaceous large volcanic eruptions are also believed to have contributed to the Cretaceous–Paleogene extinction event. Approximately 50% of all genera became extinct, including all of the non-avian dinosaurs.

Compared to the vigorous convergent plate mountain-building of the late Paleozoic, Mesozoic tectonic deformation was comparatively mild. The sole major Mesozoic orogeny occurred in what is now the Arctic, creating the Innuitian orogeny, the Brooks Range, the Verkhoyansk and Cherskiy Ranges in Siberia, and the Khingan Mountains in Manchuria. This orogeny was related to the opening of the Arctic Ocean and subduction of the North China and Siberian cratons under the Pacific Ocean. Nevertheless, the era featured the dramatic rifting of the supercontinent Pangaea. Pangaea gradually split into a northern continent, Laurasia, and a southern continent, Gondwana. This created the passive continental margin that characterizes most of the Atlantic coastline (such as along the U.S. East Coast) today.

By the end of the era, the continents had rifted into nearly their present form. Laurasia became North America and Eurasia, while Gondwana split into South America, Africa, Australia, Antarctica and the Indian subcontinent, which collided with the Asian plate during the Cenozoic, the impact giving rise to the Himalayas.

The Triassic was generally dry, a trend that began in the late Carboniferous, and highly seasonal, especially in the interior of Pangaea. Low sea levels may have also exacerbated temperature extremes. With its high specific heat capacity, water acts as a temperature-stabilizing heat reservoir, and land areas near large bodies of water—especially the oceans—experience less variation in temperature. Because much of the land that constituted Pangaea was distant from the oceans, temperatures fluctuated greatly, and the interior of Pangaea probably included expansive areas of desert. Abundant red beds and evaporites such as halite support these conclusions, but evidence exists that the generally dry climate of the Triassic was punctuated by episodes of increased rainfall. Most important humid episodes were the Carnian Pluvial Event and one in the Rhaetian, few million years before the Triassic-Jurassic extinction event.

Sea levels began to rise during the Jurassic, which was probably caused by an increase in seafloor spreading. The formation of new crust beneath the surface displaced ocean waters by as much as 200 m (656 ft) more than today, which flooded coastal areas. Furthermore, Pangaea began to rift into smaller divisions, bringing more land area in contact with the ocean by forming the Tethys Sea. Temperatures continued to increase and began to stabilize. Humidity also increased with the proximity of water, and deserts retreated.

The climate of the Cretaceous is less certain and more widely disputed. Higher levels of carbon dioxide in the atmosphere are thought to have caused the world temperature gradient from north to south to become almost flat: temperatures were about the same across the planet. Average temperatures were also higher than today by about 10°C. In fact, by the middle Cretaceous, equatorial ocean waters (perhaps as warm as 20°C in the deep ocean) may have been too warm for sea life, and land areas near the equator may have been deserts despite their proximity to water. The circulation of oxygen to the deep ocean may also have been disrupted. For this reason, large volumes of organic matter that was unable to decompose accumulated, eventually being deposited as “black shale”.

Not all of the data support these hypotheses, however. Even with the overall warmth, temperature fluctuations should have been sufficient for the presence of polar ice caps and glaciers, but there is no evidence of either. Quantitative models have also been unable to recreate the flatness of the Cretaceous temperature gradient.

Different studies have come to different conclusions about the amount of oxygen in the atmosphere during different parts of the Mesozoic, with some concluding oxygen levels were lower than the current level (about 21%) throughout the Mesozoic, some concluding they were lower in the Triassic and part of the Jurassic but higher in the Cretaceous, and some concluding they were higher throughout most or all of the Triassic, Jurassic and Cretaceous.

The dominant land plant species of the time were gymnosperms, which are vascular, cone-bearing, non-flowering plants such as conifers that produce seeds without a coating. This is opposed to the earth’s current flora, in which the dominant land plants in terms of number of species are angiosperms. One particular plant genus, Ginkgo, is thought to have evolved at this time and is represented today by a single species, Ginko biloba. As well, the extant genus Sequoia is believed to have evolved in the Mesozoic.

The extinction of nearly all animal species at the end of the Permian Period allowed for the radiation of many new lifeforms. In particular, the extinction of the large herbivorous pareiasaurs and carnivorous gorgonopsians left those ecological niches empty. Some were filled by the surviving cynodonts and dicynodonts, the latter of which subsequently became extinct. Some plant species had distributions that were markedly different from succeeding periods; for example, the Schizeales, a fern order, were skewed to the Northern Hemisphere in the Mesozoic, but are now better represented in the Southern Hemisphere.

Recent research indicates that the specialized animals that formed complex ecosystems, with high biodiversity, complex food webs and a variety of niches, took much longer to reestablish, recovery did not begin until the start of the mid-Triassic, 4M to 6M years after the extinction and was not complete until 30M years after the Permian-Triassic extinction event. Animal life was then dominated by large archosaurian reptiles: dinosaurs, pterosaurs, and aquatic reptiles such as ichthyosaurs, plesiosaurs, and mosasaurs.

The climatic changes of the late Jurassic and Cretaceous provided for further adaptive radiation. The Jurassic was the height of archosaur diversity, and the first birds and eutherian mammals also appeared. Angiosperms radiated sometime in the early Cretaceous, first in the tropics, but the even temperature gradient allowed them to spread toward the poles throughout the period. By the end of the Cretaceous, angiosperms dominated tree floras in many areas, although some evidence suggests that biomass was still dominated by cycad and ferns until after the Cretaceous–Paleogene extinction.

Some have argued that insects diversified with angiosperms because insect anatomy, especially the mouth parts, seems particularly well-suited for flowering plants. However, all major insect mouth parts preceded angiosperms and insect diversification actually slowed when they arrived, so their anatomy originally must have been suited for some other purpose.

As the temperatures in the seas increased, the larger animals of the early Mesozoic gradually began to disappear while smaller animals of all kinds, including lizards, snakes, and perhaps primates, evolved. The Cretaceous–Paleogene extinction event exacerbated this trend. The large archosaurs became extinct, while birds and mammals thrived, as they do today.

In North America, the era began with deep sedimentary basins along the eastern, southeastern, and western sides of the continent, while the interior was dry land. As the era proceeded, the marginal seas periodically washed over the stable interior, leaving sedimentary deposits to mark their incursions. During the early part of the era, the area of exposed Precambrian, or shield, rocks in central Canada were eroding, supplying sediment to the basins from the interior. Beginning in the Ordovician Period, mountain building intermittently proceeded in the eastern part of the Appalachian region throughout the rest of the era, bringing in new sediments. Sediments washing from the Acadian Mountains filled the western part of the Appalachian basins to form the famous coal swamps of the Carboniferous Period. In North America carboniferous is not generally used instead the time is divided between Mississippian and Pennsylvanian period because of differences in the sedimentary rock deposited in this time. The Mississippian is characterized by limey sediments deposited in shallow seas, typically with abundant crinoidal fossils as in the Burlington fm. The Pennsylvanian typically is characterized by terrestrial sediments such as sands, shale and most importantly coal. Most of our oil and gas are obtained from Pennsylvanian sediments. Where this has been stripped, as in the Ozark domal region, oil is not typically available.

Paleoclimatic studies and evidence of glaciers indicate that central Africa was most likely in the polar regions during the early Paleozoic. During the early Paleozoic, the huge continent Gondwanaland had either formed or was forming. By mid-Paleozoic, the collision of North America and Europe produced the Acadian-Caledonian uplifts, and a subduction plate uplifted eastern Australia. By the late Paleozoic, continental collisions formed the supercontinent Pangaea and resulted in some of the great mountain chains, including the Appalachians, Urals, and mountains of Tasmania.

Geologically, the Paleozoic starts shortly after the breakup of a supercontinent called Pannotia. Throughout the early Paleozoic, the Earth’s landmass was broken up into a substantial number of continents. Towards the end of the era, the continents gathered together into a supercontinent called Pangaea, which included most of the Earth’s land area.

The Ordovician and Silurian periods were warm greenhouse periods, with the highest sea levels of the Paleozoic (200 m above today’s); the warm climate was interrupted only by a 30 million years cool period, the Early Palaeozoic Icehouse, culminating in the Hirnantian glaciation.

The early Cambrian climate was probably moderate at first, becoming warmer over the course of the Cambrian, as the second-greatest sustained sea level rise in the Phanerozoic got underway. However, as if to offset this trend, Gondwana moved south with considerable speed, so that, in Ordovician time, most of West Gondwana (Africa and South America) lay directly over the South Pole. The early Paleozoic climate was also strongly zonal, with the result that the “climate”, in an abstract sense became warmer, but the living space of most organisms of the time—the continental shelf marine environment—became steadily colder. However, Baltica (Northern Europe and Russia) and Laurentia (eastern North America and Greenland) remained in the tropical zone, while China and Australia lay in waters which were at least temperate. The Early Paleozoic ended, rather abruptly, with the short, but apparently severe, late Ordovician ice age. This cold spell caused the second-greatest mass extinction of Phanerozoic time. Over time, the warmer weather moved into the Paleozoic Era.

The middle Paleozoic was a time of considerable stability. Sea levels had dropped coincident with the ice age, but slowly recovered over the course of the Silurian and Devonian. The slow merger of Baltica and Laurentia, and the northward movement of bits and pieces of Gondwana created numerous new regions of relatively warm, shallow sea floor. As plants took hold on the continental margins, oxygen levels increased and carbon dioxide dropped, although much less dramatically. The north–south temperature gradient also seems to have moderated, or metazoan life simply became hardier, or both. At any event, the far southern continental margins of Antarctica and West Gondwana became increasingly less barren. The Devonian ended with a series of turnover pulses which killed off much of Middle Paleozoic vertebrate life, without noticeably reducing species diversity overall.

The late Paleozoic was a time which has left us a good many unanswered questions. The Mississippian began with a spike in atmospheric oxygen, while carbon dioxide plummeted to unheard-of lows. This destabilized the climate and led to one, and perhaps two, ice ages during the Carboniferous. These were far more severe than the brief Late Ordovician Ice; but, this time, the effects on world biota were inconsequential. By the Cisuralian, both oxygen and carbon dioxide had recovered to more normal levels. On the other hand, the assembly of Pangaea created huge arid inland areas subject to temperature extremes. The Lopingian is associated with falling sea levels, increased carbon dioxide and general climatic deterioration, culminating in the devastation of the Permian extinction.

While macroscopic plant life appeared early in the Paleozoic and possibly late in the Neoproterozoic, it mostly remained aquatic until sometime in the Silurian and Devonian, when it began to transition onto dry land. Terrestrial flora reached its climax in the Carboniferous, when towering lycopsid rainforests dominated the tropical belt of Euramerica. Climate change caused the Carboniferous Rainforest Collapse which fragmented this habitat, diminishing the diversity of plant life in the late Carboniferous and Permian.

A noteworthy feature of Paleozoic life is the sudden appearance of nearly all of the invertebrate animal phyla in great abundance at the beginning of the Cambrian. The first vertebrates appeared in the form of primitive fish, which greatly diversified in the Silurian and Devonian. The first animals to venture onto dry land were the arthropods. Some fish had lungs and strong, bony fins and could crawl onto the land also. The bones in their fins eventually evolved into legs and they became the first tetrapods. Amphibians were the dominant tetrapods until the mid-Carboniferous, when climate change greatly reduced their diversity. Later, reptiles prospered and continued to increase in number and variety by the late Permian.

The geologic record of the Proterozoic is much better than that for the preceding Archean. In contrast to the deep-water deposits of the Archean, the Proterozoic features many strata that were laid down in extensive shallow epicontinental seas; furthermore, many of these rocks are less metamorphosed than Archean-age ones, and plenty are unaltered. Study of these rocks shows that the eon continued the massive continental accretion that had begun late in the Archean, as well as featured the first definitive supercontinent cycles and wholly modern orogenic activity

The first known glaciations occurred during the Proterozoic; one began shortly after the beginning of the eon, while there were at least four during the Neoproterozoic, climaxing with the Snowball Earth of the Sturtian and Marinoan glaciations.

One of the most important events of the Proterozoic was the accumulation of oxygen in the Earth’s atmosphere. Though oxygen is believed to have been released by photosynthesis as far back as Archean times, it could not build up to any significant degree until chemical sinks — unoxidized sulfur and iron — had been filled; until roughly 2.3 billion years ago, oxygen was probably only 1% to 2% of its current level. Banded iron formations, which provide most of the world’s iron ore, were also a prominent chemical sink; their accumulation ceased after 1.9 billion years ago, either due to an increase in oxygen or a more thorough mixing of the oceanic water column.

Red beds, which are colored by hematite, indicate an increase in atmospheric oxygen after 2 billion years ago; they are not found in older rocks. The oxygen buildup was probably due to two factors: a filling of the chemical sinks, and an increase in carbon burial, which sequestered organic compounds that would have otherwise been oxidized by the atmosphere.

Throughout the history of the Earth, there have been times when the continental mass came together to form a supercontinent, followed by the break-up of the supercontinent and new continents moving apart again. This repetition of tectonic events is called a Wilson cycle. It is at least clear that, about 1,000–830 Ma, most continental mass was united in the supercontinent Rodinia. Rodinia was not the first supercontinent; it formed at about 1.0 Ga by accretion and collision of fragments produced by breakup of the older supercontinent, called Nuna or Columbia, which was assembled by global-scale 2.0–1.8 Ga collisional events. This means plate tectonic processes similar to today’s must have been active during the Proterozoic.

After the break-up of Rodinia about 800 Ma, it is possible the continents joined again around 550 Ma. The hypothetical supercontinent is sometimes referred to as Pannotia or Vendia. The evidence for it is a phase of continental collision known as the Pan-African orogeny, which joined the continental masses of current-day Africa, South America, Antarctica and Australia. It is extremely likely, however, that the aggregation of continental masses was not completed, since a continent called Laurentia (roughly equivalent to current-day North America) had already started breaking off around 610 Ma. It is at least certain that by the end of the Proterozoic eon, most of the continental mass lay united in a position around the south pole.

The first advanced single-celled, eukaryotes and multi-cellular life, Francevillian Group Fossils, roughly

The blossoming of eukaryotes such as acritarchs did not preclude the expansion of cyanobacteria; in fact, stromatolites reached their greatest abundance and diversity during the Proterozoic, peaking roughly 1200 million years ago.

Classically, the boundary between the Proterozoic and the Phanerozoic eons was set at the base of the Cambrian period when the first fossils of animals including trilobites and archeocyathids appeared. In the second half of the 20th century, a number of fossil forms have been found in Proterozoic rocks, but the upper boundary of the Proterozoic has remained fixed at the base of the Cambrian, which is currently placed at 542 Ma.